“[W]ith the gene itself steering evolution, the fact that the drone shared its DNA with the queen meant that its servitude guarantees not the individual’s survival, but the endurance of the genes they share. Or as the Anglo-Indian biologist JBS Haldane put it: “Would I lay down my life to save my brother? No, but I would to save two brothers or eight cousins.””

— Adam Rutherford, The Selfish Gene turns 40 (The Guardian, May 29, 2016)